Rivers and streams, where they are
unpolluted, are frequently host to a rich variety of mosses and liverworts.
Very few species are able to grow permanently submerged, but many are characteristic
of the flood zone of rivers, growing on rocks or tree bases. A few are restricted
to this habitat. Accumulations of silt at the banks of rivers, rich in nutrients,
can also carry a distinctive bryophyte flora.
Streams in shaded woodland gullies, especially where there is some limestone,
are also often rich in bryophytes.
Some of the mosses and liverworts that are found in Welsh rivers and streams
are listed below.
Mosses
- Amblystegium riparium
- This is a common moss on wood, rocks and stones at or near the waters
edge in slow-moving rivers and streams, canals, ditches and ponds. It
forms loose matts flattened against the substrate, and with the leaves
widely-spreading from the stem. The leaves are up to 4mm in length,
tapering to fine points, and with a nerve running for about three-quarters
their length.
- Brachythecium rivulare
- A common species in and by rocky streams. It has a glossy dark-green
appearance, with the shoot tips often conspicuously lighter in colour.
The leaves have a nerve runnig for about three-quarters their length,
and distinct auricles.
- Cinclidotus fontinaloides
- This is a robust moss which bears a superficial resemblance to
Fontinalis species (hence fontinaloides). Unlike those species,
however, its leaves are distinctly bordered and have a nerve which runs
the entire length. It grows in the flood zone of rivers.
- Cryphaea lamyana
- A rare species found on rocks and trees in the flood zone of rocky
rivers. A fairly robust species with shoots having a cylindrical appearance,
and often curved. The leaves are ovate with a nerve running about three-quarters
their length. Cells are rounded. This is a Red
Data Book species in Britain, and in Wales is only known from a
section of the River Teifi. It is specially protected by inclusion on
Schedule 8 of the Wildlife and Countryside
Act.
- Dicranella palustris
- This is a very distinctive moss with its upright stems and spreading,
translucent leaves. It is a characteristic moss of mountains streams
where there is a slow trickle of water.
- Fontinalis antipyretica
- This is one of the few Welsh mosses which normally grows submerged
in water. It is a robust species, dark-green in colour and with the
leaves arranged on the stems in three ranks. The leaves are folded sharply
in two equal halves, giving the appearance of a small boat. This is
mainly a plant of slow-flowing rivers and streams. (The closely related
Fontinalis squamosa has concave leaves, not sharply folded, and
is found in fast-flowing waters).
- Hyocomium armoricum
- This densely-branched moss occurs abundantly in upland streams in
Wales, forming carpets which are bright golden-green in colour, or darker
with bright-green shoot tips. It is able to tolerate fast flowing water
and can often be found by waterfalls.
- Leskea polycarpa
- This species occurs in very similar situations to the previous one,
and is common. It is a slender plant, and has tiny leaves up to 1mm
in length. The leaves are rounded in their lower half, then drawn to
a pointed tip. Unusually amongst pleurocarpous mosses the leaf
cells are rounded or hexagonal.
- Myrinia pulvinata
- This is another species associated with silt in the flood zone of
rivers. It occurs only on tree boles an roots, however, and is much
less common than the previous two species (it is listed as British Red
Data Book species). . A slender plant, occurring in soft, dull-green
patches with prostrate main stems and ascending branches.
- Rhynchostegium riparioides
- A robust species which grows on rocks low on the banks of streams
or on rocks in the streams. The stems are often long and with few branches.
Lower down they are often bare of leaves. The leaves are ovate, with
fine teeth at their margin, and have a nerve running for three-quarters
their length. The leaves tend to stand away from the stem. A common
species.
- Thamnobryum alopecurum
- This is a common moss on shaded rocks in many streams and rivers in
Wales (though not where the water is very acid). It is a robust, branched
species, with a bright- to dark-green, almost blackish colour. Its stems
are noticeably wiry. The manner of the branching (with the branches
clustered toward the tips of the stems) gives the plant a tree-like
(dendroid) appearance.
- Tortula latifolia
- This moss grows on silted tree bases and rocks and walls by the side
of slow-flowing rivers and streams. It is usually confined to a zone
subject to flooding, but not permanently submerged. It has leaves which
are broadest above their middle, and with a nerve running the entire
length (but not extending beyond the leaf tip as in most other species
of Tortula). It is most common in the eastern half of Wales.
Liverworts
- Anthelia julacea
- This species forms dense tufts or wide cushions by mountains streams,
dull bluish green when wet, and pale gray when dry. It is a slender
plant, with leaves deeply divided and arranged in three ranks (the underleaves
in this species are similar to the laterals). In the UK it is mainly
confined to North Wales, North-West England and Scotland.
- Chiloscyphus polyanthos
- A leafy liverwort which forms loose prostrate mats of flattened shoots,
with oblong, undivided leaves arranged neatly on either sideof the stem.
Underleaves are present and divided into two sharp prongs. It grows
on wet rocks or soil by streams and rivers.
- Conocephalum conicum
- This robust thallose liverwort grows on basic or neutral soil beside
streams in wooded valleys, as well as other damp, shaded habitats. Its
surface is marked with roughly hexagonal divisions, at the centre of
each is a small pore, visible with the naked eye. When crushed the thallus
gives off a distinctive scent.
- Jubula hutchinsiae
- A plant of very wet, shaded rocks in wooded ravines, especially around
waterfalls and cascades, occurring as dark-green patches. It has leaves
which are bilobed, with the very much smaller lobe helmet-shaped and
the larger lobe coarsely toothed. Bi-lobed underleaves are present.
In the UK it is calssified as Nationally Scarce; in Wales, it is most
common in the North.
- Porella pinnata
- This is another plant which grows in the flood zone of rivers, on
rocks, and less usually on tree-boles and roots. It is a fairly robust
leafy liverwort, with leaves crowded and overlapping. The leaves are
divided into two lobes, but with one lobe very much smaller than the
other, lying underneath the larger, and adpressed to the stem. Oval-shaped
underleaves are also present. This species is Nationally Rare
in the UK, and is mainly confined to Wales and SW England, but may be
locally abundant.
- Riccia huebeneriana
- A small thallose liverwort with a rosette-like habit. It is a colonist
of recently exposed mud at the edge of ponds lakes and reservoirs, as
well as rivers. It is a Red Data Book species in the UK and known from
only a few sites in Wales. Where it does occur it can be in some quantity
for relatively short periods of time (perhaps weeks), and then disappear
again.
- Scapania undulata
- This is a very common leafy liverwort of upland streams (and also
occurs in a wide variety of other wet habitats). In common with other
Scapania species, its leaves are divided into two lobes, with
the smaller folded back over the larger. The leaf margins are usually
finely-toothed.
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