Bogs
are extremely important habitats for mosses, which are often the dominant
plants. Some of Wales' rarest mosses and liverworts are found in these habitats.
The word "bog" is a colloquial
term for what is more precisely referred to as an ombrotrophic mire
A mire is a wetland
that supports a vegetation which is normally peat-forming.
An ombrotrophic mire
is so called because its water supply, and also its nutrients, are derived
directly from rainfall (or snow, hail etc.) alone. (Gk. ombros
= a storm of rain, trophos = feeder).
This contrasts with fens,
which are also peatlands, but which receive at least some of their water
supply and nutrients from the mineral groundwater table of the surrounding
landscape and are thus termed geotrophic (earth-fed).
Bogs, because they receive
nutrients only from precipitation, are oligotrophic (poorly fed).
As a result, they are also often very acidic environments. Indeed, ombrotrophic
bogs represent one of the most nutrient-poor and acidic environments in
the Welsh natural landscape.
There are various types of
bogs or ombrotrophic mires, but two of the most important are raised
bogs and blanket bogs.
Raised bogs are so called
because of the domed profile of the peat - highest at the centre of the
bog and gently sloping away at the edges, like an inverted saucer. Often
they are formed over shallow lakes which became infilled with vegetation
- climatic conditions then resulted in plant remains accumulating as peat
eventually forming the bog as it exists today. The surface of a raised
bog is not usually flat, but is often made up of a series of "hummocks"
and "hollows".
One of the largest raised
bogs in Wales is Cors Caron near Tregaron.
Blanket bogs are more typical
of upland areas in Wales, and their shape much more reflects the contours
of the landscape - overlaying them like a blanket!
Several sub-types of both
raised and blanket bogs have been described. More information can be found
in: Lindsay, R. Bogs: The Ecology, Classification and Conservation
of Ombrotrophic Mires (Scottish Natural Heritage, 1995).
Some of the
mosses and liverworts to be found in bogs in Wales are listed below:
Mosses
One group of mosses in particular
are associated with bogs - these are the Sphagnum mosses. They
are of particular importance because of their moisture-holding properties,
and because some of them are major contributors to the accumulation of
peat. (More information on Sphagnum
mosses ).
- Sphagnum
austinii
- This moss has the typical
structure of Sphagnum mosses generally. It is distinguished by the rather
short and stubby branches, rather like little cigars, and its predominantly
golden-brown colour,intermixed with some green and yellow. It was once
a common species of raised bogs in Wales, and a major hummock-forming
species. It has suffered a steady decline since the Middle Ages, for
reasons which are not understood, and is now rare on raised bogs in
Wales. In the UK it is classified as Nationally Scarce. Climate
change, drainage of raised mires, and burning have all been suggested
as possible causes of this decline.
- Sphagnum capillifolium
- This is one of the Sphagnum
species associated with drier parts of bogs - the raised hummocks that
occur in many bogs, for example. It varies in colour from green through
to a deep wine-red, and grows in compact cushions.
- Sphagnum cuspidatum
- This is a species particularly
associated with bog pools and flooded peat cuttings, where it grows
submerged in water. In general appearance when wet it has been likened
to a "drowned kitten". Green in colour, its branch leaves are notably
long and narrow.
- Sphagnum
fuscum
- This species is a rich
brown in colour and grows in very compact, neat cushions on the tops
of hummocks on raised bogs. It is rare in Wales, and known from only
three sites, where it occurs in small quantities. In the UK it is
Nationally Scarce.
- Sphagnum
pulchrum
- The Latin suffix "pulchrum"
means "beautiful", and indeed this is one of the most attractive of
the Sphagnum mosses, with its regularly-arranged leaves and its deep
orange-brown colour. It is exclusively a plant of raised bogs and occurs
in wet depressions or pools, sometimes replacing the more common Sphagnum
cuspidatum. Sphagnum pulchrum is classified as Nationally
Scarce in the UK as a whole, and is known from only three sites
in Wales. It can, however, be abundant where it does occur.
- Sphagnum magellanicum
- This is a robust species,
a dark red in colour. Like Sphagnum tenellum it occupies a niche
intermediate between the tops of hummocks and pools or depressions.
-
- Sphagnum tenellum
- This is a species which
occupies an intermediate niche, not in the wettest, nor in the driest,
parts of bogs. It is usually yellowish in appearance, and is characterised
by very concave branch leaves which give something of the appearance
of threaded beads, and the stem leaves which are large relative to the
branch leaves and the plant as a whole.
Liverworts
- Cephalozia connivens
- This is a slender, pale-green
to yellowish plant which grows over mosses or directly on peat. It has
leaves which are bilobed, with the lobes pointed and turned towards
each other.
- Kurzia pauciflora
- This is a tiny plant, dark-green
in colour with its leaves divided into four finger-like lobes. It grows
amongst Sphagnum mosses.
- Mylia anomala
- Superficially similar in
some respects to the previous species, and often growing closely associated
with it. This species is somewhat lighter in colour, and has very large
leaf cells - easily visible with a hand lens.
- Odontoschisma sphagni
- This plant occurs as scattered
stems or sometimes extensive wefts amongst Sphagnum mosses. It
has two rows of round leaves, and is brown in colour.
- Pallavicinia
lyellii
- This is a thallose liverwort
found (in Wales) in wet hollows between tussocks of Purple Moor-grass
on the edge of raised bogs. It is readily identified by its very prominent
midrib, which is raised from the upper surface of the thallus. It is
a UK Red Data Book species,and is currently known form only two sites
in Wales.
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